Paul S. Keim is the director of pathogen genomics at TGen and is the Cowden endowed chair in microbiology at Northern Arizona University in Flagstaff. His specialty is the development of DNA fingerprinting assays for understanding and tracking especially dangerous diseases such as anthrax and plague. During the 2001 anthrax letter attacks, Keim diverted his laboratory and personal efforts to the DNA analysis of the anthrax strain from the letters. His work resulted in one of the most tangible forensic leads in the investigation. Keim earned a doctorate in plant biochemistry from KU in 1981. He continues to assist the national intelligence community in the area of biological weapons and genomic analysis.